Haven't looked in some time - but goal was to keep same what could be. Only 3.3V signals welcome.
Compare the PJRC cards - they are more alike than different as far as the pins - though ALT/secondary options not so much maybe?
LOCKABLE is special order - nothing different if left unlocked and used normally. Once LOCKED - see the PJRC info page on that - only a properly encrypted HEX ... eHex ... can be uploaded to execute and any effort to pull the code from Flash will find encrypted bit jumble.